The story is set in the mid-21st century. Global warming has led to ecological disasters all over the world, and a drastic reduction of the human population. Mankind's efforts to maintain civilization lead to the creation of "mechas," advanced humanoid robots capable of emulating human thoughts and emotions. Among these androids there is an advanced prototype model named David, a mecha created by the Cybertronics company to resemble a human child and to "feel" love for its human owners. They test their creation on one of their employees, Henry Swinton and his wife Monica. The Swintons have a son, Martin, who has been placed in suspended animation until a cure can be found for his rare disease. Although Monica is initially frightened of David, she eventually warms to him after activating his imprinting protocol, which irreversibly causes David to feel love for her as a child loves a parent. As he continues to live with the Swintons, David is befriended by Teddy, a mecha toy, who takes upon himself the responsibility of David's well being. Martin is suddenly cured and brought home; a sibling rivalry ensues between Martin and David. Martin's scheming behavior backfires when he and his friends activate David's self-protection programming at a pool party. Martin is saved from drowning but David's actions prove too much for Henry. It is decided for David to be destroyed at the factory where he was built, but Monica rather leaves him (alongside Teddy) in a forest to live as unregistered mechas. David is captured for an anti-mecha Flesh Fair, an event where useless mechas are destroyed before cheering crowds. David is nearly killed, but the crowd is swayed by his realistic nature and he escapes, along with Gigolo Joe (Jude Law), a male prostitute mecha on the run after being framed for murder.
The two set out to find the Blue Fairy, whom David remembers from the story The Adventures of Pinocchio. As in the story, he believes that she will transform him into a real boy, so Monica will love him and take him back. Joe and David make their way to the decadent metropolis of Rouge City. Information from a holographic volumetric display answer engine called "Dr. Know" eventually leads them to the top of the Rockefeller Center in the flooded ruins of Manhattan. David's human creator, Professor Hobby, enters and excitedly tells David that finding him was a test, which has demonstrated the reality of his love and desire. A disheartened David attempts to commit suicide by falling from a ledge into the ocean, but Joe rescues him, just as he is captured by the authorities.
David and Teddy take a submersible to the fairy, which turns out to be a statue from a submerged attraction at Coney Island. Teddy and David become trapped when the park's ferris wheel falls on their vehicle. Believing the Blue Fairy to be real, he asks to be turned into a real boy, repeating his wish without end, until the ocean freezes. 2000 years later, Manhattan is buried under several hundred feet of glacial ice, and humans are extinct.[1] Mechas have evolved into an alien-looking humanoid form.[2] They find David and Teddy: functional mechas who knew living humans. David wakes up and realizes the fairy was fake. Using David's memories, the mechas reconstruct the Swinton home, and explain to him via a mecha of the Blue Fairy that he cannot become human. However, they recreate Monica from a lock of her hair which has been faithfully saved by Teddy, but she will live for only a single day and the process cannot be repeated. David spends the happiest day of his life playing with Monica and Teddy. Monica tells David that she loves him and has always loved him as she drifts to sleep for her final time. This was the "everlasting moment" he had been looking for, he closes his eyes, falls asleep for his first time, and goes "to that place where dreams are born".
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